Systematik der Agaminae

Die Unterfamilie Agaminae [GRAY, 1827]

Agaminae ist eine von sechs und gleichzeitig die namensgebende Unterfamilie der Agamen (Agamidae). Sie stammen aus Afrika, Europa und Asien. Es sind über 140 Arten aus neun Gattungen beschrieben, darunter u. a. Siedleragamen, Krötenkopfagamen und Blaukehlagamen.

Gattung Acanthocercus (Blaukehlagamen) [FITZINGER, 1843]

Acanthocercus adramitanus (Hadramautagame) [ANDERSON, 1896]
Acanthocercus annectens [BLANFORD, 1870]
Acanthocercus atricollis (Blaukehlagame) [SMITH, 1849]
Acanthocercus branchi [WAGNER, GREENBAUM & BAUER, 2012]
Acanthocercus ceriacoi [MARQUES, PARRINHA ,SANTOS, BANDEIRA, BUTLER, SOUSA & WAGNER, 2022]
Acanthocercus cuongi [NGO, LE, NGUYEN, NGUYEN, NGUYEN, PHAN, NGUYEN, ZIEGLER & DO, 2025]
Acanthocercus cyanocephalus [FALK, 1925]
Acanthocercus cyanogaster [RÜPPELL, 1835]
Acanthocercus gregorii [KLAUSEWITZ, 1957]
Acanthocercus grismeri [LE, NGUYEN, NGUYEN, ZIEGLER, DO & NGO, 2025]
Acanthocercus guentherpetersi [LARGEN & SPAWLS, 2006]
Acanthocercus kiwuensis [KLAUSEWITZ, 1957]
Acanthocercus margaritae [WAGNER, BUTLER, CERÍACO & BAUER, 2021]
Acanthocercus minutus [KLAUSEWITZ, 1957]
Acanthocercus phillipsii [BOULENGER, 1895]
Acanthocercus ugandaensis [KLAUSEWITZ, 1957]
Acanthocercus yemensis [KLAUSEWITZ, 1954]

Gattung Agama (Echte Agamen) [DAUDIN, 1802]

Agama aculeata [MERREM, 1820]
- Agama aculeata aculeata [MERREM, 1820]
- Agama aculeata distanti [BOULENGER, 1902]
Agama africana [HALLOWELL, 1844]
Agama agama (Siedleragame) [LINNAEUS, 1758]
Agama anchietae (Anchieta's Agame) [BOCAGE, 1896]
Agama armata (Bodenagame) [PETERS, 1855]
Agama atra [DAUDIN, 1802]
Agama bibronii (Atlasagame) [DUMÉRIL, 1851]
Agama bocourti [ROCHEBRUNE, 1884]
Agama boensis [MONARD, 1940]
Agama bottegi [BOULENGER, 1897]
Agama boueti (Mali Agame) [CHABANAUD, 1917]
Agama boulengeri [LATASTE, 1886]
Agama caudospinosa [MEEK, 1910]
- Agama caudospinosa caudospinosa [MEEK, 1910]
- Agama caudospinosa spawlsi [WAGNER 2010]
Agama congica (Angola-Agame) [PETERS, 1877]
Agama cristata [MOCQUARD, 1905]
Agama dodomae [LOVERIDGE, 1923]
- Agama dodomae dodomae [LOVERIDGE, 1923]
- Agama dodomae ufipae [LOVERIDGE, 1932]
Agama doriae (Nigeria Agame) [BOULENGER, 1885]
- Agama doriae benuensis [MONARD, 1951]
- Agama doriae doriae [BOULENGER, 1885]
Agama etoshae [MCLACHLAN, 1981]
Agama finchi [BÖHME, WAGNER, MALONZA, LÖTTERS & KÖHLER, 2005]
- Agama finchi finchi [BÖHME, WAGNER, MALONZA, LÖTTERS & KÖHLER, 2005]
- Agama finchi leucerythrolaema [WAGNER, FREUND, MODRY, SCHMITZ & BÖHME, 2011]
Agama gracilimembris (Benin Agame) [CHABANAUD, 1918]
Agama hartmanni (Hartmanns Agame) [PETERS, 1869]
Agama hispida (Südliche Stachelagame) [KAUP, 1827]
- Agama hispida hispida [KAUP, 1827]
- Agama hispida makarikarika [FITZSIMONS, 1932]
Agama hulbertorum [WAGNER, 2014]
Agama insularis [CHABANAUD, 1918]
Agama kaimosae [LOVERIDGE, 1935]
Agama kirkii (Kirks Agame) [BOULENGER, 1885]
- Agama kirkii fitzsimonsi [LOVERIDGE, 1950]
- Agama kirkii kirkii [BOULENGER, 1885]
Agama knobeli [BOULENGER & POWER, 1921]
Agama lanzai [WAGNER, LEACHÉ, MAZUCH & BÖHME, 2013]
Agama lebretoni [WAGNER, BAREJ & SCHMITZ, 2009]
Agama lionotus (Ostafrikanische Siedleragame) [BOULENGER, 1896]
- Agama lionotus elgonis [LÖNNBERG, 1922]
- Agama lionotus lionotus [BOULENGER, 1896]
Agama lucyae [WAGNER & BAUER, 2011]
Agama montana (Blaukopf Bergagame) [BARBOUR & LOVERIDGE, 1928]
Agama mossambica (Mozambique Agame) [PETERS, 1854]
Agama mucosoensis (Mucoso-Siedleragame) [HELLMICH, 1957]
Agama mwanzae [LOVERIDGE, 1923]
Agama parafricana [TRAPÉ, MEDIANNIKOV & TRAPÉ, 2012]
Agama paragama (Falsche Siedleragame) [GRANDISON, 1968]
Agama persimilis [PARKER, 1942]
Agama picticauda [PETERS, 1877]
Agama planiceps (Stachelagame) [PETERS, 1862]
Agama robecchii [BOULENGER, 1892]
Agama rueppelli [VAILLANT, 1882]
- Agama rueppelli occidentalis [PARKER, 1932]
- Agama rueppelli rueppelli [VAILLANT, 1882]
- Agama rueppelli septentrionalis [PARKER, 19322]
Agama sankaranica (Senegal Agame) [MOCQUARD, 1905]
Agama somalica [WAGNER, LEACHÉ, MAZUCH & BÖHME, 2013]
Agama spinosa (Wüstenagame) [GRAY, 1831]
Agama tassiliensis [GENIEZ, PADIAL & CROCHET, 2011]
Agama turuensis (Ostafrikanische Siedleragame) [LOVERIDGE, 1896]
Agama wachirai [MALONZA, SPAWLS, FINCH & BAUER, 2021]
Agama weidholzi (Gambia Agame) [WETTSTEIN, 1932]

Gattung Bufoniceps [ARNOLD, 1992]

Bufoniceps laungwalaensis [SHARMA, 1980]

Gattung Laudakia [GRAY, 1845]

Laudakia agrorensis [STOLICZKA, 1872]
Laudakia cypriaca (Zypern-Hardun) [DAAN, 1967]
Laudakia dayana [STOLICZKA, 1871]
Laudakia melanura (Schwarze Felsenagame) [BLYTH, 1854]
- Laudakia melanura lirata [BLANDORD, 1874]
- Laudakia melanura melanura [BLYTH, 1854]
- Laudakia melanura nasiri [BAIG, 1999]
Laudakia nupta (Pakistanische Felsenagame) [DE FILIPPI, 1843]

Laudakia nupta fusco [BLANFORD, 1876]
Laudakia nupta nupta [DE FILIPPI, 1843]

Laudakia nuristanica [ANDERSON & LEVITON, 1969]
Laudakia pakistanica [BAIG, 1989] (Pakistan Agame)
- Laudakia pakistanica auffenbergi [BAIG & BÖHME, 1996]
- Laudakia pakistanica khani [BAIG & BÖHME, 1996]
- Laudakia pakistanica pakistanica [BAIG, 1989]
Laudakia papenfussi [ZHAO, 1998]
Laudakia sacra [SMITH, 1935]
Laudakia stellio (Hardun) [LINNAEUS, 1758]
- Laudakia stellio daani [BEUTLER & FRÖR, 1980]
- Laudakia stellio stellio [LINNAEUS, 1758]
Laudakia tuberculata (Kashmir-Felsenagame) [GRAY, 1827]
Laudakia vulgaris (Ägyptischer Hardun) [SONNINI & LATREILLE, 1802]

Laudakia vulgaris brachydactyla [HAAS, 1951]
Laudakia vulgaris picea (Schwarzer Hardun) [PARKER, 1935]
Laudakia vulgaris salehi [WERNER, 2006]

Laudakia vulgaris vulgaris [SONNINI & LATREILLE, 1802]
Laudakia wui [ZHAO, 1998]

Gattung Paralaudakia [BAIG, WAGNER, ANANJEVA & BÖHME, 2013]

Paralaudakia badakhshana [ANDERSON & LEVITON, 1969]
Paralaudakia bochariensis [NIKOLSKY, 1897]
Paralaudakia caucasia (Kaukasus-Agame) [EICHWALD, 1831]
Paralaudakia erythrogastra [NIKOLSKY, 1896]
Paralaudakia himalayana (Himalaya-Agame) [STEINDACHNER, 1867]
Paralaudakia lehmanni (Turkmenische Agame) [NIKOLSKY, 1896]
Paralaudakia microlepis [BLANFORD, 1874]
- Paralaudakia microlepis microlepis [BLANFORD, 1874]
- Paralaudakia microlepis taftanica [SANCHOOLI, RAHIMIAN, RASTEGAR-POUYANI & RASTEGAR-POUYANI 2015]
Paralaudakia stoliczkana [BLANFORD, 1875]
- Paralaudakia stoliczkana altaica [PETERS, 1971]
- Paralaudakia stoliczkana stoliczkana [BLANFORD, 1875]

Gattung Phrynocephalus (Krötenkopfagamen) [KAUP, 1825]

Phrynocephalus ananjevae [MELNIKOV, MELNIKOVA, NAZAROV & RAJABIZADEH, 2013]
Phrynocephalus arabicus (Arabische Krötenkopfagame) [ANDERSON, 1894]
Phrynocephalus ahvazicus [MELNIKOV, MELNIKOVA, NAZAROV, RAJABIZADEH, AL-JOHANY, AMR & ANANJEVA 2014]
Phrynocephalus alpherakii [BEDRIAGA, 1906]
Phrynocephalus axillaris [BLANFORD, 1875]
Phrynocephalus bannikovi [DAREVSKY, RUSTAMOV & SHAMMAKOV 1976]
Phrynocephalus carinipes [NIKOLSKY, 1907]
Phrynocephalus clarkorum [ANDERSON & LEVITON, 1967]
Phrynocephalus erythrurus [ZUGMAYER, 1909]
Phrynocephalus euptilopus [ALCOCK & FINN, 1897]
Phrynocephalus forsythii [ANDERSON, 1872]
Phrynocephalus frontalis (Shansi-Krötenkopfagame) [STRAUCH, 1876]
Phrynocephalus golubewii [SHENBROT & SEMYONOV, 1990]
Phrynocephalus guttatus (Gefleckter Krötenkopf) [GMELIN, 1789]
- Phrynocephalus guttatus guttatus [GMELIN, 1789]
- Phrynocephalus guttatus salsatus [GORELOV, DUNAYEV & KOTENKO, 1995]
Phrynocephalus helioscopus (Sonnengucker) [PALLAS, 1773]
- Phrynocephalus helioscopus cameranoi [BEDRIAGA 1907]
- Phrynocephalus helioscopus helioscopus [PALLAS, 1773]
- Phrynocephalus helioscopus karatauensis [SOLOVYEVA, DUNAYEV, NAZAROV & POYARKOV, 2025]
- Phrynocephalus helioscopus sergeevi [DUNAYEV, SOLOVYEVA & POYARKOV 2012]
- Phrynocephalus helioscopus turcomanus [DUNAYEV, SOLOVYEVA & POYARKOV 2012]
- Phrynocephalus helioscopus varius (Mongolischer Sonnengucker) [EICHWALD, 1831]
Phrynocephalus hispidus [BEDRIAGA, 1909]
Phrynocephalus horvathi [MEHELY 1894]
Phrynocephalus interscapularis [LICHTENSTEIN, 1856]
Phrynocephalus kangsuensis [LIANG & SHI, 2024]
Phrynocephalus kulagini [BEDRIAGA, 1909]
Phrynocephalus lutensis [KAMALI & ANDERSON, 2015]
Phrynocephalus luteoguttatus (Krötenkopfagame) [BOULENGER, 1887]
Phrynocephalus longicaudatus [HAAS, 1957]
Phrynocephalus maculatus (Gefleckte Krötenkopfagame) [ANDERSON, 1872]
Phrynocephalus melanurus [EICHWALD, 1831]
Phrynocephalus meridionalis [DUNAYEV, SOLOVYEVA & POYARKOV 2012]
Phrynocephalus moltschanowi [NIKOLSKY, 1913]
Phrynocephalus mystaceus (Bärtiger Krötenkopf) [PALLAS, 1776]
- Phrynocephalus mystaceus khorasanus [SOLOVYEVA, DUNAYEV, NAZAROV, RAJABIZADEH & POYARKOV, 2018]
- Phrynocephalus mystaceus mystaceus [PALLAS, 1776]
Phrynocephalus nejdensis [HAAS, 1957]
Phrynocephalus ornatus [BOULENGER, 1887]
Phrynocephalus parvus [TZAREVSKY 1927]
Phrynocephalus persicus [DE FILIPPI, 1863]
Phrynocephalus przewalskii [STRAUCH, 1876]
Phrynocephalus putjatai [BEDRIAGA, 1909]
Phrynocephalus raddei [BOETTGER, 1888]
- Phrynocephalus raddei boettgeri [BEDRIAGA, 1906]
- Phrynocephalus raddei raddei [BOETTGER, 1888]
Phrynocephalus reticulatus [EICHWALD, 1831]
Phrynocephalus roborowskii [BEDRIAGA, 1907]
Phrynocephalus rossikowi [NIKOLSKY, 1898]
- Phrynocephalus rossikowi rossikowi [NIKOLSKY, 1898]
- Phrynocephalus rossikowi shammakowi [SZCERBAK & GOLUBEV, 1979]
Phrynocephalus saidalievi [SATTOROV, 1981]
- Phrynocephalus saidalievi orlovae [SOLOVYEVA, DUNAYEV, NAZAROV & POYARKOV, 2025]
- Phrynocephalus saidalievi saidalievi [SATTOROV, 1981]
Phrynocephalus sakoi [MELNIKOV, MELNIKOVA, NAZAROV, AL-JOHANY & ANANJEVA, 2015]
Phrynocephalus scutellatus [OLIVIER, 1807]
Phrynocephalus sogdianus [CHERNOV 1948]
Phrynocephalus strauchi [NIKOLSKY, 1899]
Phrynocephalus theobaldi [BLYTH, 1863]
Phrynocephalus tickelii [GRAY 1845]
Phrynocephalus versicolor [STRAUCH, 1876]
- Phrynocephalus versicolor doriai [BEDRIAGA, 1909]
- Phrynocephalus versicolor siebenrocki [BEDRIAGA, 1909]
- Phrynocephalus versicolor versicolor [STRAUCH, 1876]
Phrynocephalus vindumi [GOLUBEV, 1998]
Phrynocephalus vlangalii [STRAUCH, 1876]
- Phrynocephalus vlangalii hongyuanensis [ZHAO, JIANG & HUANG, 1980]
- Phrynocephalus vlangalii lidskii [BEDRIAGA 1909]
- Phrynocephalus vlangalii vlangalii [STRAUCH, 1876]

Gattung Pseudotrapelus [FITZINGER, 1843]

Pseudotrapelus aqabensis [MELNIKOV, NAZAROV, ANANJEVA & DISI, 2012]
Pseudotrapelus chlodnickii [MELNIKOV, ŚMIEŁOWSKI, MELNIKOVA, NAZAROV & ANANJEVA, 2015]
Pseudotrapelus dhofarensis [MELNIKOV & PIERSON, 2012]
Pseudotrapelus jensvindumi [MELNIKOV, ANANJEVA & PAPENFUSS, 2013]
Pseudotrapelus neumanni [TORNIER, 1905]
Pseudotrapelus sinaitus (Sinai Agame) [HEYDEN, 1827]
- Pseudotrapelus sinaitus sinaitus [HEYDEN, 1827]
- Pseudotrapelus sinaitus werneri [MORAVEC, 2002]
Pseudotrapelus tuwaiqensis [TAMAR, UVIZL, SHOBRAK, ALMUTAIRI, BUSAIS, SALIM, ALGETHAMI, ALGETHAMI, ALANAZI, ALSUBAIE, CHIRIO, CARRANZA & ŠMÍD, 2023]

Gattung Trapelus [CUVIER, 1817]

Trapelus agilis (Steppenagame) [OLIVIER, 1807]
- Trapelus agilis agilis [OLIVIER, 1807]
- Trapelus agilis isolepis [BOULENGER, 1885]
- Trapelus agilis khuzistanensis [RASTEGAR-POUYANI, 1999]
- Trapelus agilis pakistanensis [RASTEGAR-POUYANI, 1999]
Trapelus agnetae [WERNER, 1929]
Trapelus boehmei [WAGNER, MELVILLE, WILMS & SCHMITZ, 2011]
Trapelus flavimaculatus [RÜPPELL, 1835]
Trapelus megalonyx [GÜNTHER, 1864]
Trapelus mutabilis (Wüstenagame) [MERREM, 1820]

Trapelus mutabilis mutabilis [MERREM, 1820]
Trapelus mutabilis pallidus [REUSS, 1834]
Trapelus mutabilis poppeki [WAGNER, MELVILLE, WILMS & SCHMITZ, 2011]

Trapelus persicus [BLANFORD, 1881]
- Trapelus persicus fieldii [HAAS & WERNER, 1969]
- Trapelus persicus persicus [BLANFORD, 1881]
Trapelus rubrigularis [BLANFORD, 1875]
Trapelus ruderatus (Ruinenagame) [OLIVIER, 1804]
Trapelus sanguinolentus (Steppenagame) [PALLAS, 1814]
- Trapelus sanguinolentus aralensis [LICHTENSTEIN, 1823]
- Trapelus sanguinolentus sanguinolentus [PALLAS, 1814]
Trapelus savignii [DUMÉRIL & BIBRON, 1837]
Trapelus schmitzi (Schmitz’ Agame) [WAGNER & BÖHME, 2006]
Trapelus tournevillei (Sahara Agame) [LATASTE, 1880]

Gattung Xenagama [BOULENGER, 1895]

Xenagama batillifera (Große Biberschwanzagame) [VAILLANT, 1882]
Xenagama taylori (Kleine Biberschwanzagame) [PARKER, 1935]
Xenagama wilmsi [WAGNER, MAZUCH & BAUER, 2013]
Xenagama zonura [BOULENGER, 1895]

Letzte Änderungen

2025-09-16 Phrynocephalus helioscopus karatauensis, Phrynocephalus saidalievi orlovae (ssp. nov.)

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