Systematik der Colubrinae - Teil 3
Die Unterfamilie Colubrinae (Eigentliche Nattern) [OPPEL, 1811]
Colubrinae ist eine von acht Unterfamilien der Familie Colubridae (Nattern). Als deutsche Bezeichnung werden "Eigentliche Nattern" und alternativ "Land- und Baumnattern" verwendet. Die größte Artenvielfalt befindet sich in Nordamerika, Asien, Nordafrika und dem Nahen Osten. In Europa, Südamerika, Australien und im südlichen Afrika gibt es nur wenige Arten. In Madagaskar, in der Karibik und auf den Pazifikinseln gibt es keine Vertreter der Colubrinae. Zur Unterfamilie gehören rund 900 Arten aus etwa 100 Gattungen. Aufgrund der Vielzahl der Arten haben wir die Unterfamilie in drei Kapitel unterteilt.
Gattung Rhamnophis (Schwarze Baumschlangen) [GÜNTHER, 1862]
- Rhamnophis aethiopissa [GÜNTHER, 1862]
- Rhamnophis aethiopissa aethiopissa [GÜNTHER, 1862]
- Rhamnophis aethiopissa elgonensis [LOVERIDGE, 1929]
- Rhamnophis aethiopissa ituriensis [SCHMIDT, 1923]
- Rhamnophis batesii [BOULENGER, 1908]
Gattung Rhinobothryum [WAGLER, 1830]
- Rhinobothryum bovallii [ANDERSSON, 1916]
- Rhinobothryum lentiginosum [SCOPOLI, 1785]
Gattung Rhinocheilus [BAIRD & GIRARD, 1853]
- Rhinocheilus antonii [DUGÈS, 1886]
- Rhinocheilus etheridgei (Etheridges Nasennatter) [GRISMER, 1990]
- Rhinocheilus lecontei (Nasennatter) [BAIRD & GIRARD, 1853]
Gattung Rhynchocalamus [GÜNTHER, 1864]
- Rhynchocalamus arabicus(Arabische Kukrinatter) [SCHMIDT, 1933]
- Rhynchocalamus dayanae[TAMAR, ŠMÍD, GÖÇMEN, MEIRI & CARRANZA, 2016]
- Rhynchocalamus hejazicus [LICATA, POLA, ŠMÍD, IBRAHIM, LIZ, SANTOS, PATKÓ, ABDULKAREEM, GONÇALVES, ALSHAMMARI, BUSAIS, EGAN, RAMALHO, SMITHSON & BRITO, 2024]
- Rhynchocalamus levitoni [TORKI, 2017]
- Rhynchocalamus melanocephalus (Schwarzkopf-Zwergnatter, Schwarzkopfnatter) [JAN, 1862]
- Rhynchocalamus satunini [NIKOLSKY, 1899]
Gattung Salvadora [BAIRD & GIRARD, 1853]
- Salvadora bairdi [JAN, 1860]
- Salvadora deserticola (Wüsten-Pflasternasennatter) [SCHMIDT, 1940]
- Salvadora grahamiae (Berg-Pflasternasennatter) [BAIRD & GIRARD, 1853]
- Salvadora gymnorhachis [ERNÁNDEZ-JIMÉNEZ, FLORES-VILLELA & CAMPBELL, 2019]
- Salvadora hexalepis (Westliche Pflasternasennatter) [COPE, 1867]
- Salvadora hexalepis hexalepis [COPE, 1866]
- Salvadora hexalepis klauberi [BOGERT, 1945]
- Salvadora hexalepis mojavensis [BOGERT, 1945]
- Salvadora hexalepis virgultea [BOGERT, 1936]
- Salvadora intermedia [HARTWEG, 1940]
- Salvadora lemniscata [COPE, 1895]
- Salvadora lineata [SCHMIDT, 1940]
- Salvadora mexicana [DUMÉRIL, BIBRON & DUMÉRIL, 1854]
Gattung Scaphiophis [PETERS, 1870]
- Scaphiophis albopunctatus [PETERS, 1870]
- Scaphiophis raffreyi [BOCOURT, 1875]
Gattung Scolecophis [FITZINGER, 1843]
- Scolecophis atrocinctus (Schwarzgebänderte Wühltrugnatter) [SCHLEGEL, 1837]
Gattung Senticolis [DOWLING & FRIES, 1987]
- Senticolis triaspis (Grüne Rattennatter) [COPE, 1866]
- Senticolis triaspis intermedia [BOETTGER, 1883]
- Senticolis triaspis mutabilis [COPE, 1885]
- Senticolis triaspis triaspis [COPE, 1866]
Gattung Sonora [BAIRD & GIRARD, 1853]
- Sonora aemula [COPE, 1879]
- Sonora annulata [BAIRD, 1859]
- Sonora annulata annulata [BAIRD, 1859]
- Sonora annulata klauberi [STICKEL, 1941]
- Sonora cincta [COPE, 1861]
- Sonora episcopa [KENNICOTT, 1859]
- Sonora fasciata [COPE, 1892]
- Sonora michoacanensis [DUGÈS, 1884]
- Sonora mosaueri [STICKEL, 1938]
- Sonora mutabilis [STICKEL, 1943]
- Sonora occipitalis (Westliche Schaufelnasenschlange) [HALLOWELL, 1854]
- Sonora palarostris (Sonora-Schaufelnasenschlange) [KLAUBER, 1937]
- Sonora palarostris organica [KLAUBER, 1951]
- Sonora palarostris palarostris [KLAUBER, 1937]
- Sonora punctatissima [VAN DENBURGH & SLEVIN, 1921]
- Sonora savagei [CLIFF, 1954]
- Sonora semiannulata [BAIRD & GIRARD, 1853]
- Sonora straminea (Gebänderte Sandschlange) [COPE, 1860]
- Sonora taylori [BOULENGER, 1894]
Gattung Spalerosophis [JAN, 1865]
- Spalerosophis arenarius [FISCHER, 1885]
- Spalerosophis atriceps (Diademnatter) [FISCHER, 1885]
- Spalerosophis diadema (Pakistanische Diademnatter) [SCHLEGEL, 1837]
- Spalerosophis diadema cliffordii [SCHLEGEL, 1837]
- Spalerosophis diadema diadema [SCHLEGEL, 1837]
- Spalerosophis diadema schirasianus [JAN, 1863]
- Spalerosophis dolichospilus [WERNER, 1923]
- Spalerosophis josephscorteccii [LANZA, 1964]
- Spalerosophis microlepis [JAN, 1865]
Gattung Spilotes (Hühnerfresser) [WAGLER, 1830]
- Spilotes megalolepis [GÜNTHER, 1865]
- Spilotes pullatus (Hühnerfresser) [LINNAEUS, 1758]
- Spilotes sulphureus [WAGLER, 1824]
- Spilotes sulphureus dieperinkii [SCHLEGEL, 1837]
- Spilotes sulphureus sulphureus [WAGLER, 1824]
Gattung Stegonotus [DUMÉRIL, DUMÉRIL & BIBRON, 1854]
- Stegonotus admiraltiensis [RUANE, RICHARDS, MCVAY, TJATURADI, KREY & AUSTIN, 2017]
- Stegonotus aplini [O’SHEA & RICHARDS, 2021]
- Stegonotus aruensis [DORIA, 1875]
- Stegonotus australis [GÜNTHER, 1872]
- Stegonotus ayamaru [KAISER, O’SHEA & KAISER, 2019]
- Stegonotus batjanensis [GÜNTHER, 1865]
- Stegonotus borneensis [INGER, 1967]
- Stegonotus caligocephalus [KAISER, LAPIN, O’SHEA & KAISER, 2020]
- Stegonotus cucullatus [DUMÉRIL, DUMÉRIL & BIBRON, 1854]
- Stegonotus derooijae [RUANE, RICHARDS, MCVAY, TJATURADI, KREY & AUSTIN, 2017]
- Stegonotus diehli [LINDHOLM, 1905]
- Stegonotus dorsalis[WERNER 1924]
- Stegonotus florensis [DE ROOIJ, 1917]
- Stegonotus guentheri [BOULENGER, 1895]
- Stegonotus heterurus [BOULENGER, 1893]
- Stegonotus iridis [RUANE, RICHARDS, MCVAY, TJATURADI, KREY & AUSTIN, 2017]
- Stegonotus keyensis [DORIA, 1874]
- Stegonotus lividus [DUMÉRIL, BIBRON & DUMÉRIL, 1854]
- Stegonotus melanolabiatus [RUANE, RICHARDS, MCVAY, TJATURADI, KREY & AUSTIN, 2017]
- Stegonotus modestus [SCHLEGEL, 1837]
- Stegonotus muelleri [DUMÉRIL, BIBRON & DUMÉRIL, 1854]
- Stegonotus nancuro [KAISER, KAISER, MECKE & O'SHEA, 2021]
- Stegonotus parvus [MEYER, 1874]
- Stegonotus poechi [WERNER, 1924]
- Stegonotus reticulatus [BOULENGER, 1895]
- Stegonotus sutteri [FORCART, 1953]
Gattung Stenorrhina [DUMÉRIL, 1853]
- Stenorrhina degenhardtii (Degenhardts Skorpionfresser) [BERTHOLD, 1846]
- Stenorrhina degenhardtii degenhardtii [BERTHOLD, 1846]
- Stenorrhina degenhardtii mexicana [STEINDACHNER, 1867]
- Stenorrhina degenhardtii ocellata [JAN, 1876]
- Stenorrhina freminvillei (Skorpionsfresser) [DUMÉRIL, BIBRON & DUMÉRIL, 1854]
Gattung Tantilla [BAIRD & GIRARD, 1853]
- Tantilla albiceps [BARBOUR, 1925]
- Tantilla alticola [BOULENGER, 1903]
- Tantilla andinista [WILSON & MENA, 1980]
- Tantilla armillata (Schwarznacken-Schwarzkopfschlange) [COPE, 1876]
- Tantilla atriceps [GÜNTHER, 1895]
- Tantilla bairdi [STUART, 1941]
- Tantilla berguidoi [BATISTA, MEBERT, LOTZKAT & WILSON, 2016]
- Tantilla bocourti [GÜNTHER, 1895]
- Tantilla boipiranga [SAWAYA & SAZIMA, 2003]
- Tantilla brevicauda [MERTENS, 1952]
- Tantilla briggsi [SAVITZKY & SMITH, 1971]
- Tantilla calamarina [COPE, 1866]
- Tantilla capistrata [COPE, 1876]
- Tantilla carolina [PALACIOS-AGUILAR, FUCSKO, JIMÉNEZ-ARCOS, WILSON & MATA-SILVA, 2022]
- Tantilla cascadae [WILSON & MEYER, 1981]
- Tantilla ceboruca [CANSECO-MÁRQUEZ, SMITH, PONCE-CAMPOS, FLORES-VILLELA & CAMPBELL, 2007]
- Tantilla coronadoi [HARTWEG, 1944]
- Tantilla coronata (Gekrönte Schwarzkopfschlange) [BAIRD & GIRARD, 1853]
- Tantilla cucullata [MINTON, 1956]
- Tantilla cuniculator [SMITH, 1939]
- Tantilla deppei [BOCOURT, 1883]
- Tantilla excelsa [MCCRANIEL & SMITH, 2017]
- Tantilla flavilineata [SMITH & BURGER, 1950]
- Tantilla gottei [MCCRANIEL & SMITH, 2017]
- Tantilla gracilis (Schlanke Schwarzkopfschlange) [BAIRD & GIRARD, 1853]
- Tantilla hendersoni [STAFFORD, 2004]
- Tantilla hobartsmithi [TAYLOR, 1936]
- Tantilla impensa [CAMPBELL, 1998]
- Tantilla insulamontana [WILSON & MENA, 1980]
- Tantilla jani [GÜNTHER, 1895]
- Tantilla johnsoni [WILSON, VAUGHN & DIXON, 1999]
- Tantilla lempira [WILSON & MENA, 1980]
- Tantilla lydia [ANTÚNEZ-FONSECA, CASTRO, ESPAÑA, TOWNSEND & WILSON, 2020]
- Tantilla melanocephala (Dunkelköpfige Schwarzkopfschlange) [LINNAEUS, 1758]
- Tantilla miyatai [WILSON & KNIGHT, 1987]
- Tantilla moesta [GÜNTHER, 1863]
- Tantilla nigra [BOULENGER, 1914]
- Tantilla nigriceps (Flachland-Schwarzkopfschlange) [KENNICOTT, 1860]
- Tantilla oaxacae [WILSON & MEYER, 1971]
- Tantilla olympia [TOWNSEND, WILSON, MEDINA-FLORES & HERRERA-B, 2013]
- Tantilla oolitica [TELFORD, 1966]
- Tantilla palamala [ESQUEDA, ROJAS-RUNJAIC, CORREA, ORTÍZ, GUERRERO, JIMÉNEZ, BAZÓ, MORENO-PÉREZ, AGUILAR & URRA, 2025]
- Tantilla petersi [WILSON, 1979]
- Tantilla planiceps (Westliche Schwarzkopfschlange) [BLAINVILLE, 1835]
- Tantilla psittaca [MCCRANIE, 2011]
- Tantilla relicta [TELFORD, 1966]
- Tantilla relicta neilli [TELFORD, 1966]
- Tantilla relicta pamlica [TELFORD, 1966]
- Tantilla relicta relicta [TELFORD, 1966]
- Tantilla reticulata [COPE, 1860]
- Tantilla robusta [CANSECO-MÁRQUEZ, MENDELSOHN & GUTIÉRREZ-MAYÉN, 2002]
- Tantilla rubra (Große Schwarzkopfschlange) [COPE, 1876]
- Tantilla ruficeps [COPE, 1894]
- Tantilla schistosa [BOCOURT, 1883]
- Tantilla selmae [AZEVEDO, FRANCO, MENEZES, SARAIVA-KUNZ & GRAZZIOTIN , 2024]
- Tantilla semicincta [DUMÉRIL, BIBRON & DUMÉRIL, 1854]
- Tantilla sertula [WILSON & CAMPBELL, 2000]
- Tantilla shawi [TAYLOR, 1949]
- Tantilla slavensi [PÉREZ-HIGAREDA, SMITH & SMITH, 1985]
- Tantilla stenigrammi [MCCRANIEL & SMITH, 2017]
- Tantilla striata [DUNN, 1928]
- Tantilla supracincta [PETERS, 1863]
- Tantilla taeniata [BOCOURT, 1883]
- Tantilla tayrae [WILSON, 1983]
- Tantilla tecta [CAMPBELL & SMITH, 1997]
- Tantilla tjiasmantoi [KOCH & VENEGAS, 2016]
- Tantilla trilineata [PETERS, 1880]
- Tantilla triseriata [SMITH & SMITH, 1951]
- Tantilla tritaeniata [SMITH & WILLIAMS, 1966]
- Tantilla vermiformis (Wurm-Schwarzkopfschlange) [HALLOWELL, 1861]
- Tantilla vulcani [CAMPBELL, 1998]
- Tantilla wilcoxi [STEJNEGER, 1902]
- Tantilla yaquia [SMITH, 1942]
Gattung Tantillita [H. M. SMITH, 1941]
- Tantillita brevissima [TAYLOR, 1937]
- Tantillita canula [COPE, 1876]
- Tantillita lintoni [SMITH, 1940]
- Tantillita lintoni lintoni [SMITH, 1940]
- Tantillita lintoni rozellae [PEREZ-HIGAREDA, 1985]
Gattung Telescopus [WAGLER, 1830]
- Telescopus beetzi [BARBOUR, 1922]
- Telescopus crocheti [RIBEIRO-JÚNIOR, KOCH, FLECKS, CAMPBELL, CALVO, SPAWLS, VIDAL & MEIRI, 2025]
- Telescopus dhara (Israelische Katzennatter) [FORSKAL, 1775]
- Telescopus fallax (Europäische Katzennatter) [FLEISCHMANN, 1831]
- Telescopus fallax cyprianus [BARBOUR & AMARAL, 1927]
- Telescopus fallax fallax [FLEISCHMANN, 1831]
- Telescopus fallax iberus [EICHWALD, 1831]
- Telescopus fallax pallidus [STEPANEK, 1944]
- Telescopus fallax syriacus [BOETTGER, 1889]
- Telescopus finkeldeyi (Damara-Katzennatter) [HAACKE, 2013]
- Telescopus forskali [RIBEIRO-JÚNIOR, KOCH, FLECKS, CAMPBELL, CALVO, SPAWLS, VIDAL & MEIRI, 2025]
- Telescopus geniezi [RIBEIRO-JÚNIOR, KOCH, FLECKS, CAMPBELL, CALVO, SPAWLS, VIDAL & MEIRI, 2025]
- Telescopus gezirae (Sudan-Katzennatter) [BROADLEY, 1994]
- Telescopus guentheri [ANDERSON, 1895]
- Telescopus hoogstraali (Sinai-Katzennatter) [SCHMIDT & MARX, 1956]
- Telescopus insularis [RIBEIRO-JÚNIOR, KOCH, FLECKS, CAMPBELL, CALVO, SPAWLS, VIDAL & MEIRI, 2025]
- Telescopus mazuchi [RIBEIRO-JÚNIOR, KOCH, FLECKS, CAMPBELL, CALVO, SPAWLS, VIDAL & MEIRI, 2025]
- Telescopus nigriceps (Schwarzkopf-Katzennatter) [AHL, 1924]
- Telescopus obtusus (Ägyptische Katzennatter) [REUSS, 1834]
- Telescopus pulcher (Somalische Katzennatter) [SCORTECCI, 1935]
- Telescopus reussi [RIBEIRO-JÚNIOR, KOCH, FLECKS, CAMPBELL, CALVO, SPAWLS, VIDAL & MEIRI, 2025]
- Telescopus rhinopoma (Persische Katzennatter) [BLANFORD, 1874]
- Telescopus salvadori [RIBEIRO-JÚNIOR, KOCH, FLECKS, CAMPBELL, CALVO, SPAWLS, VIDAL & MEIRI, 2025]
- Telescopus semiannulatus (Südafrikanische Tigernatter) [SMITH, 1849]
- Telescopus semiannulatus polystictus [MERTENS, 1954]
- Telescopus semiannulatus semiannulatus [SMITH, 1849]
- Telescopus smidi [RIBEIRO-JÚNIOR, KOCH, FLECKS, CAMPBELL, CALVO, SPAWLS, VIDAL & MEIRI, 2025]
- Telescopus somalicus [PARKER, 1949]
- Telescopus tessellatus (Irakische Katzennatter) [WALL, 1908]
- Telescopus tripolitanus (Nordafrikanische Katzennatter) [WERNER, 1909]
- Telescopus urii [RIBEIRO-JÚNIOR, KOCH, FLECKS, CAMPBELL, CALVO, SPAWLS, VIDAL & MEIRI, 2025]
- Telescopus variegatus (Westafrikanische Katzennatter) [REINHARDT, 1843]
- Telescopus wangariae [RIBEIRO-JÚNIOR, KOCH, FLECKS, CAMPBELL, CALVO, SPAWLS, VIDAL & MEIRI, 2025]
Gattung Thelotornis [A. SMITH, 1849]
- Thelotornis capensis [SMITH, 1849]
- Thelotornis kirtlandii (Lianennatter, Graue Baumnatter) [HALLOWELL, 1844]
- Thelotornis mossambicanus [BOCAGE, 1895]
- Thelotornis oatesi [GÜNTHER, 1881]
- Thelotornis uluguruensis [BOADLEY & WALLACH, 2002]
- Thelotornis usambaricus [BROADLEY, 2001]
Gattung Thrasops [HALLOWELL, 1857]
- Thrasops flavigularis [HALLOWELL, 1852]
- Thrasops jacksonii (Schwarze Baumschlange) [GÜNTHER, 1895]
- Thrasops occidentalis [PARKER, 1940]
- Thrasops schmidti (Schmidts Schwarze Baumschlange) [LOVERIDGE, 1936]
Gattung Toxicodryas [HALLOWELL, 1857]
- Toxicodryas adamantea [REENBAUM, ALLEN, VAUGHAN, PAUWELS, WALLACH, KUSAMBA, MUNINGA, ARIS-TOTE, MALI, BADJEDJEA, PENNER, RÖDEL, RIVERA, STERKHOVA, JOHNSON, TAPONDJOU & BROWN, 2021]
- Toxicodryas blandingii [HALLOWELL, 1844]
- Toxicodryas pulverulenta [FISCHER, 1856]
- Toxicodryas vexator [REENBAUM, ALLEN, VAUGHAN, PAUWELS, WALLACH, KUSAMBA, MUNINGA, ARIS-TOTE, MALI, BADJEDJEA, PENNER, RÖDEL, RIVERA, STERKHOVA, JOHNSON, TAPONDJOU & BROWN, 2021]
Gattung Trimorphodon [COPE, 1861]
- Trimorphodon biscutatus (Küsten-Lyraschlange) [DUMÉRIL, BIBRON & DUMÉRIL, 1854]
- Trimorphodon lambda (Sonora-Lyraschlange) [COPE, 1886]
- Trimorphodon lyrophanes (Baja California-Lyraschlange) [COPE, 1860]
- Trimorphodon paucimaculatus [TAYLOR, 1938]
- Trimorphodon quadruplex [SMITH, 1941]
- Trimorphodon tau (Mexikanische Lyraschlange) [COPE, 1870]
- Trimorphodon tau latifascia [PETERS, 1869]
- Trimorphodon tau tau [COPE, 1870]
- Trimorphodon vilkinsonii (Texas-Lyraschlange) [COPE, 1886]
Gattung Wallaceophis [MIRZA, VYAS, PATEL, MAHETA & SANAP, 2016]
- Wallaceophis gujaratensis [MIRZA, VYAS, PATEL, MAHETA & SANAP, 2016]
Gattung Xenelaphis [GÜNTHER, 1864]
- Xenelaphis ellipsifer (Gefleckte Braunnatter) [BOULENGER, 1900]
- Xenelaphis hexagonotus (Braunnatter) [CANTOR, 1847]
Gattung Yakacoatl [CISNEROS-BERNAL, PALACIOS-AGUILAR, HERNÁNDEZ-JIMÉNEZ, SMITH, FLORES-VILLELA, HERNÁNDEZ-MORALES, LOYOLA, CAMPILLO-GARCÍA, 2025]
- Yakacoatl tlalli [CISNEROS-BERNAL, PALACIOS-AGUILAR, HERNÁNDEZ-JIMÉNEZ, SMITH, FLORES-VILLELA, HERNÁNDEZ-MORALES, LOYOLA, CAMPILLO-GARCÍA, 2025]
Gattung Zamenis [WAGLER, 1830]
- Zamenis hohenackeri (Transkaukasische Kletternatter) [STRAUCH, 1873]
- Zamenis hohenackeri hohenackeri [STRAUCH, 1873]
- Zamenis hohenackeri lyciensis (Lykische Kletternatter) [HOFMANN, MEBERT, SCHULZ, HELFENBERGER, GÖÇMEN & BÖHME, 2018]
- Zamenis hohenackeri tauricus (Taurische Kletternatter) [WERNER, 1898]
- Zamenis lineatus [CAMERANO, 1891]
- Zamenis longissimus (Äskulapnatter) [LAURENTI, 1768]
- Zamenis persicus (Persische Kletternatter) [WERNER, 1913]
- Zamenis scalaris (Treppennatter) [SCHINZ, 1822]
- Zamenis situla (Leopardnatter) [LINNAEUS, 1758]
Letzte Änderungen
- 2026-02-17 Yakacoatl tlalli (gen. nov. & sp. nov.)
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